What is the difference between the English (Ireland) and English (Ireland Euro) site cultures?

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The difference is in the use of periods and commas for thousand and decimal separators.
 
 
For example:
 
One thousand two hundred and fifty-nine and ninety-nine cents.
 
 
English (Ireland) = 1,259.99 € (the comma is used as a thousands separator and the period as a decimal separator).

English (Ireland Euro) = 1.259,99 € (the period is used as a thousands separator and the comma as a decimal separator).